Denisovano
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 Replica de un molar denisovan.
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| Homo denisovensis o Homo altaiensis |
Humanes de Denisova o denisovanes[1] es un specie extincte del genere Homo; illes se trovava in Asia durante le Paleolithico (inter 450 milles e 40 milles annos retro).[2] Le denisovanes esseva primemente identificate in 2010 per ADN mitochondrial de un osso digital juvenil trovate in le Caverna Denisova in Siberia. Le denisovanes coexisteva con Homo sapiens moderne durante milles de annos, e alicun mesticiage occurreva. Homo heidelbergensis es typicamente considerate como le ancestre commun de Homo neanderthalensis e le Denisovanes.
Phylogenia
[modificar | modificar fonte]Un grande diversitate genetic (fr) se observa intra Denisovanes.[3][4]
Le analyse del ADN nuclear de spécimenes fossile de Homo sapiens, Neandertalianes e Denisovanes, effectuate in 2016 per le equipa del Instituto Max-Planck de anthropologia evolutionnista (Leipzig) ducite per Svante Pääbo, monstra que Neandertalianes e Denisovanes habeva un ancestre commun circa 450 ka retro, e que un altere con Homo sapiens un ancêtre commun circa 660 ka retro. Iste conclusiones resulta del analyse de duo specimenes fossile del Sima de los Huesos (es) (Espania), datate de 430 ka, (Neandertalianes pauc post la separation de Denisovanes.[5]
NB. Un studio publicate in 2020 indica que le chromosomas Y de Denisovanes se separeva circa 700 ka retro de un lineage commun al Neandertalianes analysate et humanes moderne; iste duo ultimates se diversificava 37 ka AP. Iste phylogenia es non conforme con isto revelate per le genoma autosomal (en) (non sexual) et revela un hybridation de Neanderthalianes recente con un lineage precoce de Homo sapiens qui conduce a reimplaciar le chromosoma Y (heritate directemente del patre) o le ADN mitochondrial (heritate del matre) de iste individuos per material genetic de H. sapiens.[6]
Le analyse comparative del genomas fossile e del introgressiones denisovan intra le genomas oceanian actual, conduce a distinger Denisovianes del nord (fossiles del Altai o chinese) e del sud (sin fossile cognite)[7] :
| ancestre commun (ca 650 ka AP) |
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Specimenes
[modificar | modificar fonte]| Name | Fossile | Datation | Discoperta | Loco | Individuo | Publication | Imagine | GenBank |
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| Denisova 3 (also known as X Woman)[8][9][10] |
phalange (en) distal (5e digito) | 76.2–51.6 ka[11] | 2008 | cava Denisova (Russia) | femina de 13.5 annos | 2010 |  |
NC013993 |
| Denisova 4[8][12] | 2e o 3e molar superior permanente | 84.1–55.2 ka[11] | 2000 | cava Denisova (Russia) | adulto mascule | 2010 | |
FR695060 |
| Denisova 8[13] | 3e molar superior permanente | 136.4–105.6 ka[11] | 2010 | cava Denisova (Russia) | adulto mascule | 2015 | KT780370 | |
| Denisova 2[14] | 2e molar inferior caduc | 194.4–122.7 ka[11] | 1984 | cava Denisova (Russia) | femina adolescente | 2017 | KX663333 | |
| Xiahe mandibula[15] | mandibula partial | > 160 ka | 1980 | cava Baishiya (China) | 2019 | 
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| Denisova 11 (fr) (dicite "Denny", hybride Denisovan x Neanderthal)[16] |
fragmento ossee de brazo o gambo | 118.1–79.3 ka[11] | 2012 | cava Denisova (Russia) | femina de 13 annos | 2016 | 
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| Denisova 13[17] | fragmento de osso parietal | trovate in strato 22[17] de ~285±39 ka[4] | 2019 | cava Denisova (Russia) | attendite | |||
| TNH2-1[18] | 1e o 2e molar inferior permanente | 164–131 ka | 2018 | cava Tam Ngu Hao 2 (Laos) | puera de 3.5 ad 8.5 annos | 2022 | 
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Un molar denisovan -
Le distribution e evolution del denisovanos -
Objectos del Caverna Denisova -
Objectos del Caverna Denisova
Vide etiam
[modificar | modificar fonte]Referentias
[modificar fonte]- ↑ Derivation (in ordine alphabetic): (ca) Denissovans || (de) Denisova-Mensch || (en) Denisovan || (es) Hombre de Denísova || (fr) Homme de Denisova || (it) Homo di Denisova || (pt) Hominídeo de Denisova || (ro) Omul de Denisova || (ru) Денисовский человек
- ↑ Molar hallado sugiere que el hombre de denísova también prospero en los trópicos
- ↑ M. Meyer, M. Kircher, M.-T. Gansauge et al. (2012-08-30). "A High-Coverage Genome Sequence from an Archaic Denisovan Individual". Science. doi:..
- ↑ 4,0 4,1 Guy S. Jacobs, Georgi Hudjashov, Lauri Saag, Pradiptajati Kusuma, Chelzie C. Darusallam, Daniel J. Lawson, Mayukh Mondal, Luca Pagani, François-Xavier Ricaut, Mark Stoneking, Mait Metspalu, Herawati Sudoyo, J. Stephen Lansing & Murray P. Cox (2019). "Multiple Deeply Divergent Denisovan Ancestries in Papuans". Cell 177 (4): 1010-1021.e32. doi:. ISSN 0092-8674.
- ↑ M. Meyer, J.-L. Arsuaga, C. de Filippo et al. (2016). "Nuclear DNA sequences from the Middle Pleistocene Sima de los Huesos hominins". Nature 531: 504-7. doi:.
- ↑ Martin Petr et al. (2020-03-09). "The evolutionary history of Neandertal and Denisovan Y chromosomes". bioRxiv. doi:.
- ↑ Pour la Science, décembre 2019, p. 32.
- ↑ 8,0 8,1 "Genetic history of an archaic hominin group from Denisova Cave in Siberia" (2010). Nature 468 (7327): 1053–60. doi:. PMID 21179161. PMC:4306417. Bibcode: 2010Natur.468.1053R.
- ↑ "Morphology of the Denisovan phalanx closer to modern humans than to Neanderthals" (2019). Science Advances 5. doi:. PMID 31517046. Bibcode: 2019SciA....5.3950B.
- ↑ "The complete mitochondrial DNA genome of an unknown hominin from southern Siberia" (in en) (2010). Nature 464 (7290): 894–897. doi:. ISSN 1476-4687. PMID 20336068. Bibcode: 2010Natur.464..894K.
- ↑ 11,0 11,1 11,2 11,3 11,4 "Age estimates for hominin fossils and the onset of the Upper Palaeolithic at Denisova Cave" (2019). Nature 565 (7741): 640–644. doi:. PMID 30700871. Bibcode: 2019Natur.565..640D. Archived 6 maio 2020 at the Wayback Machine
- ↑ "Fossil genome reveals ancestral link" (2010). Nature 468 (7327): 1012. doi:. PMID 21179140. Bibcode: 2010Natur.468.1012C.
- ↑ "Nuclear and mitochondrial DNA sequences from two Denisovan individuals" (2015). Proceedings of the National Academy of Sciences 112 (51): 15696–700. doi:. PMID 26630009. Bibcode: 2015PNAS..11215696S.
- ↑ "A fourth Denisovan individual" (2017). Science Advances 3 (7): e1700186. doi:. PMID 28695206. Bibcode: 2017SciA....3E0186S.
- ↑ "A late Middle Pleistocene Denisovan mandible from the Tibetan Plateau" (2019). Nature 569 (7756): 409–412. doi:. PMID 31043746. Bibcode: 2019Natur.569..409C.
- ↑ "Identification of a new hominin bone from Denisova Cave, Siberia using collagen fingerprinting and mitochondrial DNA analysis" (2016). Scientific Reports 6. doi:. PMID 27020421. Bibcode: 2016NatSR...623559B.
- ↑ 17,0 17,1 "A parietal fragment from Denisova cave" (2019). 88th Annual Meeting of the American Association of Physical Anthropologists. Archived 26 septembre 2019 at the Wayback Machine
- ↑ "A Middle Pleistocene Denisovan molar from the Annamite Chain of northern Laos" (2022). Nature Communications 13 (2557). doi:. PMID 35581187. Bibcode: 2022NatCo..13.2557D.
